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What does it mean by "PC" in this article?

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Answer from @user438383, @matteo-ferla and @m, converted from comments [anyone, feel free to edit this answer to improve it]:

According to the intro it means 'pseudocenter'. All measurements are relative to an assigned point because a protein doesn't have a 'centre' they assigned it as a pseudocentre.

Note that PC normally means principal component, but that does not seem to be the case here.

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